A revision of Timonius (Rubiaceae) in Kinabalu Park, Borneo, with notes on typification and species distinction

A comprehensive revision of the genus Timonius (Rubiaceae) in Kinabalu Park, Borneo, identifies 13 species, including seven likely endemic to the Kinabalu massif—four of which are restricted to ultramafic substrates. The study designates two lectotypes and proposes two new synonyms, addressing long-standing taxonomic ambiguities. Researchers examined specimens from multiple herbaria (BO, BRUN, K, L, SAN, SING, SNP) using traditional herbarium taxonomy methods, providing critical clarity for this understudied genus in West Malesia.

Chen, J., Tan, H., & Wong, K. M. (2015). A revision of Timonius (Rubiaceae) in Kinabalu Park, Borneo, with notes on typification and species distinction. Plant Ecology and Evolution, 148(3), 420–430. https://doi.org/10.5091/plecevo.2015.1073